Africa's Lake Tanganyika has heated up sharply over the past 90 years and is now warmer than at any time for at least 1,500 years, a scientific paper has warned, adding that fish and wildlife are threatened. The lake, which straddles the border between Tanzania in East Africa and the Democratic Republic of the Congo, is the world's second largest by volume and its second deepest. Lead scientist on the project Jessica Tierney said the sharp rise in temperature coincided with rises in human emissions of greenhouse gases seen in the past century, so the study added to evidence that emissions are warming the planet. The 'Great Lakes' such as Tanganyika, Malawi and Kenya's Lake Turkana were formed millions of years ago by the tectonic plate movements that tore Africa's Great Rift Valley. Some 10 million people live around Tanganyika and depend upon it for drinking water and food, mostly fish. Geologists at Rhode Island's Brown University used carbon dating to measure the age of sediments on the lake floor. They then tested fossilised micro-organisms whose membranes differ at various temperatures to gauge how hot it was at times past. The results were published in Nature Geoscience. Most climate change studies have focused on the atmosphere, but increasingly scientists are studying the effects on the oceans, seas and lakes, which all absorb a huge amount of heat. The paper argues that recent rises in temperature are correlated with a loss of biological productivity in the lake, suggesting higher temperatures may be killing life. "Lake Tanganyika has become warmer, increasingly stratified and less productive over the past 90 years," the paper says. "Unprecedented temperatures and a ... decrease in productivity can be attributed to (human) ... global warming." The rise in temperature over the past 90 years was about 0.9 degrees Celsius and was accompanied by a drop in algae volumes.